The Cycad Pages
Cycas undulata

Desf. ex Gaudich., Voy. Uranie, Bot.: 434 (1826); Desf.,: 29 (1829), nom. nud. C. circinalis [subsp. circinalis] forma undulata (hort. ex Gaudich.) Schuster, Pflanzenr. 99: 66 (1932) [note that Schuster did not include the rank of subspecies, although technically required to do so]. C. rumphii forma undulata (Desf. ex Gaudich.) Kanehira, J. Jap. Bot. 14(9): 587 (1938).

Type: hort. Paris, 1820, ex herb. Houllet (holotype: P!).

The name is attributed to Desfontaines (1829), who had published it as a nomen nudum based on material annotated by Gaudichaud-Beaupré and held in P. This specimen (above) is clearly of an immature leaf of C. circinalis from plants of unknown origin cultivated at Paris. The name must therefore fall into the synonymy of Cycas circinalis

Although described very early in the history of the genus, this name has been poorly understood throughout its history. It was formally named in 1829 by French explorer and botanist C. Gaudichaud-Beaupre (1829), in his treatment of the botany in M.L. de Freycinet's account of voyage of the ---- (Gaudichaud 1829). He ascribed the name to Renee Louiche Desfontaines(). Desfontaines had published it (without a description) earlier in 1829 in a listing of plants in the king's collection, although he in turn had taken the name from a herbarium specimen that had been annotated by Gaudichaud. Gaudichaud noted that the country of origin was unknown, and that it had been long cultivated in the king's garden.

Both Miquel (1861, 1868) and de Candolle (1868) recorded it as an uncertain species, Miquel first suggesting that it may have been the same as C. glauca, and later suggesting synonymy with C. rumphii. De Candolle proposed C. thouarsii at a possible synonym. The name C. undulata did not arise in the literature again until Schuster (1932) assigned it to material from The Marianne group, as C. circinalis forma undulata. He had also, however, included C. tonkinensis and C. bellefontii as synonyms, although both were recorded as having come from Tonkin. Kanehira (1938)Kanehira (1938) mistakenly followed Schuster, but correctly recognised the affinities of the Micronesian cycads in transferring Schuster's forma undulata to C. rumphii.

The Cycad Pages

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