|Photo Ken Hill
|The Cycad Pages
- Zamia standleyi Schutzman, Syst. Bot. 14(2): 214-219 (1989).
- "TYPE: Honduras, Atlantida, Lanatilla Valley near Tela, Aug 1984, B. Schutzman 449 (holo FLAS, iso ENA, FTG)."
|Photo Ken Hill
Named in honour of Paul Standley for his excellent thorough and prodigious
work on the flora of Mexico and Central America.
Long-lanceolate leaflets with spreading evenly
spaced marginal teeth and bearing the impression, in the form of a
slight to pronounced fold, of the adjacent leaflets as a result of
the imbricate pytxis and compression of emerging leaves and leaflets.
Additionally, the seed cones have a distinctive curved, long-acuminate
tip. The most similar species in terms of these leaflet characters
is Z. poeppigiana. Zamia poeppigiana has leaflets with
parallel margins and a pronounced falcate base, 10-15 leaves per plant
that are 1-3 m long with 20-40 pairs of leaflets, and adult plants
with trunks to 4 m tall. In contrast, Z. standleyi has upwardly
subfalcate leaflet margins and a straight attenuate base, 1-5 leaves
per plant that are less than 1 meter long with 10-15 pairs of leaflets,
and adult plants with subterranean stems.
Distribution and habitat:
Zamia standleyi is endemic to Honduras where it occurs from sea
level to 200 m in semi-xeric woodlands to secondary scrub vegetation
and even at the margins of cultivated gardens and fields.
Zamia standleyi seems to do well even in disturbed situations.
Numerous cones and seeds are produced in habitat and in cultivation.
However, caution should be exercised in the case of assuming viable
seed production in this species. A female plant growing at the New
York Botanical Garden has produced from 1-2 cones with mature seeds
every year for the past 17 years. Yet no male plants or other pollen
producing plants were present and pollination did not occur.
Accordingly these "seeds" contain no embryos. This
is the only known case in Zamia of the failure of ovules
to abort in the absence of pollination. Thus for estimates of
viable seed production in the field, seedlings need to be observed.
1997 IUCN Red List of Threatened Plants
|Photo Dennis Stevenson
|Photo Ken Hill
Stem subterranean and tuberous, 5-12 cm diam.
Cataphylls of two forms either triangular basally with
stipules and acuminate apically, 3-6 cm long, 1.2-2.8 cm wide;
or when subtending cones elongate triangular without stipules,
5-12 cm long, .75 -1.5 cm wide.
Leaves 1-5, slightly to strongly recurved, oblong,
0.2-1 m long; petiole terete, to 60 cm long, sparsely to heavily
armed with prickles; rachis terete, rarely with a few prickles,
to 45 cm long, 10-15 pairs of leaflets.
Leaflets subcoriaceous to coriaceous, long-lanceolate,
apex acute to attenuate, margins with distinct spreading and
evenly spaced teeth in the upper third, the median ones 20-45
cm long, 1-4 cm wide.
Pollen cones 1-3, decumbent, cylindrical, light-brown
to brown, 6-10 cm long, 1-2 cm in diameter; peduncle 2-4 cm long.
Seed cones usually solitary, cylindrical to slightly
ovoid, light-brown to brown, long-acuminate apically with the acumen
curved, 8-12 cm long, 3-8 cm in diameter; peduncle 2.5-4 cm long.
Seeds red, ovoid, 3 cm long, 2 cm in diameter.
2n = 16.