Type genus: Eucalyptus L'Herit., Sert. Angl.
Leaves disjunct-opposite or rarely opposite. Hairs absent, or less often unicellular (usually thick-walled) hairs radiating from raised glands on juvenile plants. Oil ducts absent from pith and petiole. Crystalliferous fibres present. Paratracheal parenchyma scanty, not confluent or banded. Renantherin present or absent, macrantherin absent. Inflorescence with fixed opposite phyllotaxy, flowers stalked or sessile, usually 4-merous, stone cells absent from sepals and petals. Stamens numerous, not bundled in mature flowers; stamens all fertile. Ovary 3-5(-6)-carpellate; ovules anatropous, arranged in distinct vertical rows on the placenta. Outer integument of ovule with crystalline inner epidermis; seed testa derived from 2 integuments.