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Family Fabaceae - faboideae

Description:

Family FABACEAE

Trees, shrubs or herbs, sometimes climbing or trailing, occasionally spiny.

Leaves alternate or sometimes opposite or whorled, usually compound (variously bipinnate, pinnate or palmate) or simple, sometimes reduced to scales or modified into phyllodes; stipules usually present and sometimes spinose, stipels often present.

Inflorescences racemose or paniculate, sometimes umbellate or cymose or flowers solitary; bracts and bracteoles commonly below flowers, often caducous. Flowers actinomorphic or zygomorphic, usually bisexual, commonly 5-merous. Sepals usually 5, free or more often variously fused into a toothed tube. Petals usually 5, free or sometimes fused, equal or unequal. Stamens commonly 10 or sometimes <10 or numerous, free or variously fused, sometimes reduced to staminodes; anthers 2-locular, opening by longitudinal slits or rarely by pores; uniform or dimorphic. Ovary superior; carpel solitary; ovules usually 2–many, attached to the upper suture; style and stigma terminal.

Fruit usually a pod (legume), mostly dehiscent along both sutures or only along upper suture or sometimes indehiscent or rarely a lomentum [or drupaceous or nut-like]; seeds 1 to many, often with a hard impervious coat, aril often developed but often inconspicuous.

Distribution and occurrence: World: c. 700 genera, 18000 species, widespread, especially tropical to temperate regions. Australia: c. 180 genera, 1900 species, all States.

Fabaceae: Family Relationships and phylogeny
(from the Angiosperm Phylogeny Site)

The family is of great economic importance for food, timber, forage crops, dyes and horticulturally. Most species have root nodules containing bacteria (Rhizobium species) which are capable of taking up atmospheric nitrogen and converting it into other nitrogenous compounds. Thus many species are capable of growing well in relatively poor soils, and any are commonly grown as fertilizer crops. Following recent trends this family is accepted with 3 subfamilies (often treated as 3 separate families) distinguished as follows.

 Key to the family FABACEAE 
1Flowers zygomorphic; petals imbricate, often unequal; stamens 10 or fewer, not forming the conspicuous part of the inflorescence; leaves various, mostly simple or pinnately compound, only occasionally bipinnately compound or reduced to phyllodes.2
2Flowers slightly zygomorphic; corolla not papilionaceous, petals ± equal, free, upper petal usually borne internally to the adjacent lateral petals; stamens 10 or fewer, free.Fabaceae - Caesalpinioideae
2*Flowers strongly zygomorphic; corolla papilionaceous, petals not equal, ± united; upper petal (standard) borne externally to the adjacent lateral petals (wings), lower petals ± fused (keel); stamens 10, often united into a tube, or the uppermost filament free, or less commonly all free.Fabaceae - Faboideae
1*Flowers actinomorphic; petals valvate, equal; stamens usually numerous, forming the conspicuous part of the inflorescence; leaves often bipinnately compound or reduced to phyllodes.Fabaceae - Mimosoideae

Description of subfamily: Trees, shrubs or herbs, sometimes climbing or twining.

Leaves simple or compound (mostly pinnate or palmate) with 1-many leaflets, rarely absent; stipules usually present.

Inflorescences various; each flower on a pedicel subtended by a lower bract and with often 2 similar bracteoles just below the calyx or attached on the calyx tube. Flowers strongly zygomorphic. Calyx mostly unequally toothed, often 2-lipped. Corolla papilionaceous, petals unequal; upper petal (the standard) usually largest and borne externally to the adjacent lateral petals (wings), lower petals innermost and mostly fused (the keel) and enfolding the stamens and gynoecium. Stamens 10, free or more often monadelphous with the filaments fused to form a closed or open sheath around the gynoecium, or the uppermost filament often ± free so that the stamens are diadelphous; anthers 2-locular, dehiscing by longitudinal slits, all uniform or sometimes dimorphic and alternating in size.

Fruit commonly a pod or sometimes a lomentum; seeds usually arillate, though often inconspicuous.


Distribution and occurrence: World: c. 500 gen., 12 000 spp., widespread, chiefly trop. to temp. regions. Aust.: c. 150 gen., 1100 spp., all States.

External links:
Wikipedia

The seeds and pods of many of the herbaceous species are eaten by humans as food; these are especially valued for their high protein content. Some, especially species of Trifolium and Medicago, are cultivated and used for feeding to livestock or ploughed in as a fertilizer. A number of species are used for timber, while many species are widely grown as ornamentals. This subfamily is often treated as the family Fabaceae s.str. or Papilionaceae. Bituminaria, Centrosema, Lablab, Phaseolus, Spartium are included in the key to genera (marked by **). Species of these genera are cultivated as ornamentals or food plants; however, they are not known to be fully naturalized in N.S.W., as a result they are not described in the text.

 Key to the genera 
1All leaves with 1 leaflet, simple, reduced, or absent.2
Leaves compound, comprised of 2 or more leaflets, rarely some leaves 1-foliolate.4
2Leaves simple or 1-foliolate.3
Leaves absent or reduced to spines or scales.
                       Back to 1
7
3Leaves simple.17
Leaves 1-foliolate, an articulation at junction of petiole with lamina.
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41
4Leaflets odd in number, rachis ending in a leaflet or character not obvious.5
Leaflets even in number, rachis often ending in a bristle or tendril.
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50
5Leaflets 3.6
Leaflets 5 or more (or more rarely some leaves 3-foliolate).
                       Back to 4
97
6Leaves palmately 3-foliolate, all 3 petiolules ± equal.56
Leaves pinnately 3-foliolate, central petiolule longer than lateral petiolules.
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67
7Stamens all free.8
Stamens variously fused.
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12
8Flowers yellow, in long racemes and panicles; calyx teeth equal, shorter than tube; pods ovoid, thin, indehiscent.Viminaria
Not as above.
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9
9Flowers in axillary racemes, sometimes umbellate or reduced to 1 flower; barren bracts present on peduncle; pods ± triangular or (rarely) ovoid.Daviesia
Combination of characters not as above.
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10
10Upper 2 calyx teeth much enlarged and mostly fused; bracteoles large, round, caducous; pods globose.Sphaerolobium
Combination of characters not as above.
                       Back to 9
11
11Calyx teeth imbricate in bud.Isotropis
Calyx teeth valvate in bud.
                       Back to 10
Jacksonia
12Upper stamen ± free, lower 9 joined in a sheath.13
All 10 stamens fused in a tube or sheath.
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15
13Trees with red to orange flowers 3–5 cm long; leafless at flowering.Erythrina
Shrubs or subshrubs with ± pink flowers <1 cm long; leaves sometines present at flowering.
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14
14Indumentum absent or hairs simpleCarmichaelia
Indumentum of two-armed hairs on most parts of the plant
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Indigofera
15All 10 stamens joined in a tube; prickly shrubs with yellow flowers.Ulex
All 10 stamens joined in a sheath split on upper side; plants and flower colour variable.
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16
16Anthers with perceptible connective, all dorsifixed, uniform in size; branches flat with blunt apex.Bossiaea
Anthers with an inconspicuous connective, alternately basifixed and dorsifixed the latter often smaller; branches terete or flat with rigid, sharply pointed apex.
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Templetonia
17Stipules fused behind the axillary bud, margins scarious.Pultenaea
Stipules free or absent.
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18
18Stamens free or fused at very base.19
Stamens variously fused.
                       Back to 17
32
19Upper 2 calyx teeth fused into a broad notched lip, lower 3 very small and narrow; bracteoles large, round, caducous; leaves tiny, caducous; pods globose.Sphaerolobium
Not as above.
                       Back to 18
20
20Ovary partly divided by a longitudinal partition inside and/or pods longitudinally grooved above.Mirbelia
Combination of characters not as above.
                       Back to 19
21
21Ovules 4 or more.22
Ovules 2.
                       Back to 20
27
22Standard purple-veined behind; upper 2 caly teeth fused most of the way into a broad lip.Isotropis
Combination of characters not as above.
                       Back to 21
23
23Calyx teeth abutted in bud; pods ± globose.Gompholobium
Calyx teeth overlapping in bud; pods ovoid or more elongated.
                       Back to 22
24
24Keel much shorter than wings or beaked; leaves mostly alternate.Chorizema
Keel about twice as long as wings; leaves ± opposite or whorled.
                       Back to 23
25
25Stipules ± rigid, recurved or spreading; hairs on stems laterally attached, ± 2-branched; bracts simple.26
Stipules bristly or absent; hairs on stems simple; bracts 3-lobed.
                       Back to 24
Oxylobium
26Margins of leaves recurved; leaves alternate, opposite, or whorled, upper surface often wartyCallistachys
Margins of leaves not recurved; leaves usually opposite, upper surface not usually warty
                       Back to 25
Podolobium
27Pods triangular (sometimes turgid) glabrous; flowers in often modified or reduced racemes with barren bracts on peduncle; bracteoles absent.Daviesia
Combination of characters not as above.
                       Back to 21
28
28Leaves with recurved to revolute margins; seeds without an aril.29
Leaves flat with flat to involute margins or terete to trigonous and grooved above; seeds with aril.
                       Back to 27
30
29Bracteoles absent.Aotus
Bracteoles present, 4–15 mm long, usually green and leaf-like.
                       Back to 28
Phyllota
30Leaves opposite, in 2 rows.Eutaxia
Leaves alternate, scattered or crowded.
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31
31Standard much broader than long.Dillwynia
Standard slightly longer than broad.
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Almaleea
32All 10 stamens fused from base in a tube closed or open on upper side.33
Upper stamen free at least at base, other 9 joined in a sheath.
                       Back to 18
39
33All stamens fused in a closed tube.34
All stamens fused in a sheath open along upper side.
                       Back to 32
35
34Calyx 2-lipped, not as above; anthers not bearded.Cytisus
Calyx with 5 small teeth, split to base on upper side; anthers bearded at base.
                       Back to 33
Spartium
35Calyx usually with 5 (rarely 4) subequal teeth; alternate anthers long and short with longer ones more than one and a half times longer.Crotalaria
Upper 2 calyx teeth joined higher than others or greatly enlarged; anthers uniform or alternate anthers longer but not more than one and a half times as long as shorter ones.
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36
36Anthers with perceptible connective, all dorsifixed, uniform in size; leaves distichous.37
Anthers without conspicuous connective, alternately dorsifixed and basifixed, the former often smaller; leaves spirally inserted.
                       Back to 35
38
37Pods not thickly and narrowly margined or keeled above, with flattish valves; leaves absent, alternate or opposite; upper calyx teeth not as enlarged as below.Bossiaea
Pods with a thin conspicuous wing above, valves rolling back; leaves opposite; upper calyx teeth greatly enlarged, >6 times as long as lower teeth.
                       Back to 36
Platylobium
38Petals blue or purple; pods not or scarcely longer than broad.Hovea
Petals yellow, red or reddish purple; pods at least one and a half times longer than broad.
                       Back to 36
Templetonia
39Anthers tipped by a small raised gland; covered with ± appressed, laterally attached hairs.Indigofera
Anthers not as above.
                       Back to 32
40
40Rigid shrubs with spiny branches; pods ± straight and smooth.Alhagi
More or less prostrate herbs, branches not spiny; pods twisted and ± spiny.
                       Back to 39
Scorpiurus
41Stipules fused behind the axillary bud, margins scarious.Pultenaea
Stipules free or absent.
                       Back to 3
42
42Stamens all free.Isotropis
Stamens variously fused.
                       Back to 41
43
43All 10 stamens fused, in a sheath split along upper side, at least towards base.44
Stamens all or mostly fused but not as above.
                       Back to 42
47
44Calyx with 5 (rarely 4) ± equal teeth; alternate anthers more than one and a half times longer than others.Crotalaria
Upper 2 calyx teeth joined higher than others or greatly enlarged; anthers uniform or not differing by more than one and a half times in length.
                       Back to 43
45
45Anthers with inconspicuous connective, alternately basifixed and dorsifixed, the latter often smaller; leaves spirally inserted.Templetonia
Anthers with perceptible connective, all dorsifixed, uniform in size; leaves distichous.
                       Back to 44
46
46Pods not or relatively thickly and narrowly margined or keeled above, valves flattish; upper calyx teeth not as enlarged as below.Bossiaea
Pods with thin conspicuous wing above, valves thin and rolling back; upper calyx teeth nearly as long as standard, enormously expanded relative to lower teeth.
                       Back to 45
Platylobium
47Pods transversely jointed or constricted, divided into 1–8, indehiscent, 1-seeded articles.122
Pods continuous, not jointed, often short and 1- or 2-seeded.
                       Back to 43
48
48All 10 stamens joined in a closed tube; erect shrubs.Cytisus
Upper stamen free, other 9 stamens joined in an open sheath; climbing or prostrate subshrubs.
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Hardenbergia
50Leaves not ending in a tendril or not; climbing herbs.51
Leaves not ending in a tendril; shrubs or non-climbing herbs.
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54
51Leaves not ending in a tendril; seeds glossy red and blackAbrus
Leaves ending in a tendril; seeds not as above
                       Back to 50
52
52Stipules of similar appearance and size to leaflets (or larger).Pisum
Stipules smaller and narrower than leaflets.
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53
53Leaflets one pair.Lathyrus
Leaflets 2 to many pairs.
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Vicia
54Leaflets 2–6 and >3 cm long, 10–45 mm wide; flowers >2 cm long.Vicia
Leaflets 2–60 and <3 cm long, 1–15 mm wide; flowers <1 cm long.
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55
55Leaves with >10 leaflets; erect annual, >40 cm high; fruit not jointed, not breaking into 1-seeded articles.Sesbania
Leaves with 2 or 4 leaflets; prostrate to erect subshrubs, <40 cm high; fruit jointed, breaking into 1-seeded articles.
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Zornia
56Stamens all free; leaves sessile or nearly so; pods globose or nearly so.Gompholobium
Stamens variously fused; leaves petiolate; pods various, rarely ± globose.
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57
57All 10 stamens fused in a sheath open along the upper side.58
Stamens all fused in a closed tube, or upper one wholly or partly free.
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60
58Upper 2 calyx teeth greatly enlarged and/or joined higher than lower 3.Muelleranthus
Calyx teeth ± equal, or if not equal, then not as above.
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59
59Procumbent herbs; leaves in groups of 3 to 5; pods linear to oblong.Lotononis
Not as above; pods inflated.
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Crotalaria
60Leaves dotted with glands, at least on lower surface.61
Leaves not dotted with glands.
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62
61Fruit densely glandular.Cullen
Fruit not glandular.
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Bituminaria
62Plants herbaceous (sometimes woody at base).63
Plants shrubby.
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64
63Plants twining or trailing (usually woody at base); inflorescences racemose.Glycine
Plants erect to procumbent; inflorescences head-like, spicate or umbellate.
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Trifolium
64Keel petals oblong, straight along lower surface; style curved abruptly near the apex.Genista
Keel petals curved along lower surface; style curved throughout its length.
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65
65Flowers creamy white; calyx tubular, 9–10 mm long; stems ± terete, with obscure ridges.Chamaecytisus
Flowers yellow; calyx campanulate, c. 6 mm long; stems prominently angular.
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Cytisus
67Stipels clearly and consistently present at base of each leaflet, rarely gland-like; frequently twining or climbing plants.68
Stipels absent, minute or inconsistently present.
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86
68Trees or robust woody climbers with leaflets at least 5 cm long, and large showy flowers at least 2 cm long, sometimes plants leafless at flowering.69
Not as above.
                       Back to 67
71
69Trees with red flowers; keel much shorter than standard.Erythrina
Woody climbers, flowers greenish or purple, blue or pink with yellow spot; keel equal to or longer than standard.
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70
70Leaves ± glabrous; keel longer than standard; pods with 2 wings along each margin, at first covered with irritant hairs, then ± glabrous.Mucuna
Leaves pubescent; keel ± equal to standard; pods not as above, softly villous.
                       Back to 69
Pueraria
71Styles bearded, or hairy more than halfway.72
Styles glabrous, or hairy less than halfway.
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76
72Styles abruptly divided into thin basal part and thickened upper part.73
Styles uniformly thick or thin, or tapering.
                       Back to 71
75
73Styles with thickened part (and keel) twisted through more than 360°.Phaseolus
Styles with thickened part twisted through no more than 360°, usually <180°.
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74
74Thickened part of styles bent through 90° just above the base, then curved slightly narrowed towards the apex, resembling a squarish hook.Macroptilium
Styles not as above, variously and smoothly curved.
                       Back to 73
Vigna
75Styles straight and blade-like; standard mostly purple.Lablab
Styles strongly curved near base and apex in one direction, gently curved in middle in opposite direction; standard pinkish.
                       Back to 72
Dipogon
76Leaves dotted with yellow resinous glands; pods acuminate, with oblique grooves between seeds, striped with maroon or purplish black.Cajanus
Not as above.
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77
77Herbs or shrubs, rarely twining; pods transversely jointed or constricted, separating into 1–8, indehiscent, 1-seeded articles.122
Stems long and trailing or twining (except rarely Hardenbergia); pods not as above.
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78
78Flowers solitary or clustered in leaf axils, or 1 or 2 on axillary peduncles.79
Flowers several to many in axillary racemes, sometimes clustered at apex of peduncle.
                       Back to 77
80
79Standard white or greenish yellow; bracteoles ± subulate.Macrotyloma
Standard showy, predominantly shades of red or purple; bracteoles absent.
                       Back to 78
Kennedia
80Pods with prominent longitudinal rib(s) near suture(s), or winged.81
Pods not as above.
                       Back to 78
82
81Lowermost calyx lobe much longer than others; pods linear with an acicular beak.Centrosema
Lower 3 calyx teeth equal or absent; pods oblong, not or slightly beaked.
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Canavalia
82Calyx with 5 teeth, or slightly 2-lipped by upper 2 teeth more joined than lower 3.83
Upper 2 calyx teeth joined to or almost to apex, calyx appearing 4-lobed or 2-lipped.
                       Back to 80
84
83Flowers solitary along rachis of inflorescence.Glycine
Flowers in clusters of 3 or more along rachis of inflorescence.
                       Back to 82
Neonotonia
84Seeds with aril; bracteoles absent.Kennedia
Seeds without aril; bracteoles present although often small or caducous.
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85
85Upper stamen joined with other 9 in tube, or becoming free; stipules produced downward from base, usually >15 mm long.Pueraria
Upper stamen free, other 9 fused; stipules smaller than above, not produced downward.
                       Back to 84
Galactia
86Stamens all fused by filaments into sheath open along upper side; pods dehiscent.87
Stamens not as above or if (rarely) so, then pods indehiscent.
                       Back to 67
88
87Fruit a pod with several seeds, not hooked at apexGoodia
Fruit a loment, commonly 1-seeded, hooked at apex by persistent style
                       Back to 86
Stylosanthes
Margins of leaflets toothed or crenulate, or if entire then plants dotted with immersed (not superficial) glands.
                       Back to 86
89
Margins of leaflets entire, not gland-dotted.
                       Back to 86
93
89Leaflets dotted with immersed black or shining glands (sometimes obscured by hairs).Cullen
Leaflets not gland-dotted.
                       Back to 88
90
90Pods enclosed in calyx; corolla persistent in fruit.Trifolium
Pods exceeding calyx; corolla not persistent in fruit.
                       Back to 89
91
91Pods ovoid, straight, indehiscent; flowers in slender elongated racemes.Melilotus
Not as above.
                       Back to 90
92
92Pods short, spirally coiled or rarely curved, usually indehiscent.Medicago
Pods long, linear, curved, tardily dehiscent.
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Trigonella
93Lower surface of leaflets, calyx and sometimes petals covered with superficial yellow or orange resinous gland dots (sometimes obscure).Rhynchosia
Not as above.
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94
94Calyx of 4 acuminate teeth by union of upper 2 teeth.Galactia
Calyx otherwise.
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95
95Flowers 1-several, clustered in leaf axils, or in heads or spikes; pods enclosed in calyx.Lespedeza
Inflorescence a raceme, sometimes few-flowered and subumbellate; pods exserted.
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96
96Styles bearded.Swainsona
Styles not bearded.
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Canavalia
97Stamens all free, or rarely filaments shortly fused at base.98
Stamens mostly fused by filaments.
                       Back to 5
100
98Shrubs; leaflets usually <5 mm wide, terminal one sessile; pods globose, <1 cm long; keel petals free or almost so.Gompholobium
Trees or tall shrubs; leaflets >5 mm wide; pods elongate, >3 cm long; keel petals fused.
                       Back to 97
99
99Leaves ± glabrous; inflorescences in axils of old leaves or on old wood; pods dehiscent, 3–5 cm diam., woody, turgid; flowers 3–4 cm long.Castanospermum
Leaves densely hairy; inflorescences terminal; pods ± indehiscent, <1 cm diam., prominently constricted between seeds; flowers <2.5 cm long.
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Sophora
100Anthers tipped by small raised gland; plant covered with more or less appressed laterally attached hairs.101
Not as above.
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102
101Standard glabrous; keel rostrate; fruit somewhat bilaterally flattened, partitions between seeds membranous, endocarp never spottedIndigastrum
Standard hairy; keel not rostrate; fruit globular to cylindric, partitions between seeds pithy; endocarp spotted
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Indigofera
102Herbs; leaves with 5 or 9 leaflets, upper 3 palmate at rachis summit, lowest 2 at its base often resembling stipules; actual stipules minute; 5 alternate or all filaments dilated upwards.Lotus
Not as above.
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103
103Stamens all fused in a sheath open along the upper side; pods dehiscent.104
Stamens variously fused, not as above, or if (rarely) so, then pods indehiscent and usually ornamented or winged.
                       Back to 102
106
104Leaves pinnately compound.Ptychosema
Leaves palmately compound.
                       Back to 103
105
105Pods inflated; style with a line of hairs on inner surface; calyx >11 mm long, glabrous; flowers yellow.Crotalaria
Pods not inflated, ± flat; style glabrous; calyx mostly <11 mm long; flowers usually blue or purple, occasionally yellow.
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Lupinus
106Stipules becoming enlarged and spinose; deciduous trees.Robinia
Stipules not spinose; plants not deciduous.
                       Back to 103
107
107Styles exserted from keel, incurved to hooked; wings petals greatly reduced; woody rainforest climbers.Austrosteenisia
Not as above.
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108
108Prostrate herbs with erect pseudo-umbels; flowers red, showy, c. 8 cm long; keel beaked; standard reflexed, acute, with a shining (usually) black boss.Swainsona
Not as above.
                       Back to 107
109
109Standard silky or woolly behind; leaflets with lateral veins numerous, parallel, ascending to the margin; pods linear or oblong, not woody; herbs or shrubs.Tephrosia
Not as above.
                       Back to 108
110
110Styles bearded.111
Styles glabrous, or pubescent near base.
                       Back to 109
112
111Standard shorter than keel; flowers >3 cm long; shrubs to 1 m high.Lessertia
Standard ± equal to or longer than keel; flowers usually <3 cm long; herbs or subshrubs.
                       Back to 110
Swainsona
112Leaflets dotted with black glands.113
Leaflets not dotted with glands.
                       Back to 110
115
113Leaves palmately compound with 3–7 leaflets.Cullen
Leaves pinnately compound, leaflets 5–17.
                       Back to 112
114
114Leaflets 5–11; flowers solitary or in axillary clusters towards end of stem.Psoralea
Leaflets 5–17; flowers in axillary spike-like racemes.
                       Back to 113
Glycyrrhiza
115Pods indehiscent, ± flat, thin, winged or keeled; woody climbers or somewhat shrubby.Brachypterum
Pods various, not as above; habit various.
                       Back to 112
116
116Pods with several seeds, not transversely jointed, 2-valved.117
Pods jointed and separating into 1-seeded articles, or short 1-seeded and bristly.
                       Back to 115
119
117Woody climbers in or near rainforest or weedy near human habitation; flowers purplish.118
Herbaceous weeds in drier habitats; flowers yellowish.
                       Back to 116
Astragalus
118Evergreen liana; infloresence a panicle; pod inflated, 3–5.2 cm wideAustrocallerya
Deciduous liana; infloresence a raceme; pod ompressed, 1.2–3 cm wide
                       Back to 117
Wisteria
119All or alternate filaments dilated upwards.120
Filaments not dilated upwards.
                       Back to 116
121
120Keel acute or beaked; pods 4-angled.Securigera
Keel obtuse; pods terete or compressed.
                       Back to 119
Ornithopus
121Pods 1-seeded, not jointed, bristly.Onobrychis
Pods jointed, of 2 or more 1-seeded articles, not bristly.
                       Back to 119
Aeschynomene
122Flowers in dense spike-like racemes; fruit ± enclosed by calyx; articles 2, folded back on each other.Uraria
Flowers in open racemes or clusters; fruit not enclosed by calyx; articles 1–8, not folded back on each other.
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123
123Leaves 1-foliolate; bracts filiform.124
Leaves mostly 3-foliolate, occasionally a few 1-foliolate or 5-foliolate; bracts not filiform.
                       Back to 122
125
124Pod separating into 1-seeded, turgid articlesAlysicarpus
Pod separating into 1-seeded, compressed articles
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Pleurolobus
125Branches spinescent; petioles 2–4 mm long; leaflets oblanceolate to oblong, 0.3–2.5 cm long, 1–6 mm wide.Pedleya
Branches not spinescent; petioles >4 mm long; leaflets various, 0.5–10 cm long, 5–45 mm wide.
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126
126Leaflets with a conspicuous silver stripe on upper surface.Desmodium
Leaflets uniformly green on upper surface.
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127
127Pod articles inflated; leaflets lanceolate, 4–10 cm long, 4–10 times longer than wide.Desmodiopsis
Pod articles flat; leaflets smaller and/or less elongate.
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128
128Leaflets hairy on both surfaces.Desmodium
Leaflets ± glabrous on upper surface.
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129
129Lower surface of leaflets sparsely hairy with minute, hooked hairs, often appearing glabrous; leaflets drying ± blue-black.Oxytes
Lower surface of leaflets sparsely to densely hairy with longer, appressed hairs, sometimes with hairs restricted to veins; leaflets not drying blue-black.
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130
130Stipules 5–15 mm long.Grona
Stipules 2–6 mm long.
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131
131Leaves with terminal petiolule longer than lateral ones, usually 2–4.5 mm long; leaflets lanceolate, oblong or oblanceolate.Desmodium
Leaves with terminal petiolule similar to lateral ones, ≤1 mm long; leaflets ± rhombic and truncate at the apex or rarely broad-obovate or rounded.
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Pullenia

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